Eur. J. Entomol. 106 (3): 441-450, 2009 | DOI: 10.14411/eje.2009.055

Revision of the Iteaphila setosa group (Diptera: Empididae)

1 All-Russian Institute of Plant Protection, Podbelskogo 3, Pushkin, St. Petersburg 189620, Russia; e-mail:
2 Ottawa Plant & Seed Laboratories - Entomology, Canadian Food Inspection Agency, K.W. Neatby Bldg., C.E.F., 960 Carling Ave., Ottawa, ON, Canada K1A 0C6; e-mail:

Six species are identified in the Iteaphila setosa group [I. arundela sp. n., I. caucasica sp. n., I. italica Loew, 1873, I. kubaniensis sp. n., I. merzi sp. n., I. setosa (Bezzi, 1924)]. This species group is distributed from southern England in the west, through southern Europe and northern Africa, to the Caucasus and Middle Asia (Uzbekistan).

Keywords: Iteaphila, Empidoidea, Empididae, taxonomy, new species, distribution

Received: May 6, 2009; Accepted: June 8, 2009; Published: July 24, 2009

Download citation


  1. BEZZI M. 1924: Missione del Dr. E. Festa in Cirenaica. XI. Ditteri di Cirenaica. Boll. Mus. Zool. Anat. Comp. R. Univ. Torino (N. S. no. 18) 39[1925]: 1-26
  2. CERRETTI P., TAGLIAPIETRA A., TISATO M., VANIN S., MASON F. & ZAPPAROLI M. (eds) 2003: Artropodi dell'orizzonte del faggio nell'appennino settentrionale Primo contributo. In: Conservazione Habitat Invertebrati 2. Gianluigi Arcari, Mantova, 254 pp
  3. CHANDLER P.J. (ed.) 1998: Checklists of insects of the British Isles (New Series). Part 1: Diptera (incorporating a list of Irish Diptera). Handbooks for the Identification of British Insects 12(1): 1-234
  4. CHVALA M. 1981: Classification and phylogeny of Empididae, with a presumed origin of Dolichopodidae (Diptera). Entomol. Scand. Suppl. 15: 225-236
  5. CHVALA M. 2008: Monograph of the genus Hilara Meigen (Diptera: Empididae) of the Mediterranean region. Stud. Dipterol. Suppl. 15, 138 pp
  6. COLLIN J.E. 1961: Empididae. In British Flies. Vol. 6. Cambridge University Press, Cambridge, 782 pp
  7. ENGEL E.O. 1941: 28. Empididae (partim). In Lindner E. (ed.): Die Fliegen der Palaearktischen Region IV, 4. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, pp. 193-272, pls 14-20
  8. GROOTAERT P. & CHVALA M. 1992: Monograph of the genus Platypalpus (Diptera: Empidoidea, Hybotidae) of the Mediterranean region and the Canary Islands. Acta Univ. Carol. (Biol.) 36: 3-226.
  9. JONG H. DE 1998: In search of historical biogeographic patterns in the western Mediterranean terrestrial fauna. Biol. J. Linn. Soc. 65: 99-164 Go to original source...
  10. KIRK-SPRIGGS A.H. & MCGREGOR G. 2008: Disjunctions in the Diptera fauna of the Mediterranean Province and southern Africa and a discussion of biogeographical considerations. Trans. R. Soc. S. Afr. 63: 1-21
  11. LOEW H. 1873: Systematische Beschreibung der bekannten europaeischen zweifluegeligen Insecten. Von Johann Wilhelm Meigen. Zehnter Theil oder vierter Supplementband. Beschreibungen europaeischer Dipteren. Dritter Band. H.W. Schmidt, Halle, viii + 320 pp
  12. MCALPINE J.F. 1981: Morphology and terminology - Adults. [Chapter] 2. In McAlpine J.F., Peterson B.V., Shewell G.E., Teskey H.J., Vockeroth J.R. & Wood D.M. (eds): Manual of Nearctic Diptera. Vol. 1. Agric. Canada Monogr. 27. Canadian Government Publishing Centre, Hull, pp. 9-63
  13. MELANDER A.L. 1928: Diptera, Fam. Empididae. In Wytsman P. (ed.): Genera Insectorum 185 [1927] . Louis DemetVerteneuil, Bruxelles, 434 pp
  14. MOULTON J.K. & WIEGMANN B.M. 2007: The phylogenetic relationships of flies in the superfamily Empidoidea (Insecta: Diptera). Mol. Phylogenet. Evol. 43: 701-713 Go to original source...
  15. SAIGUSA T. 2006: Homology of Wing Venation of Diptera. Unpublished handout distributed at the 6th International Congress of Dipterology, Fukuoka, Japan, 26 pp
  16. SHAMSHEV I.V. & SINCLAIR B.J. 2006: The genus Schistostoma Becker from Southern Africa, with an evaluation of its generic status (Diptera: Dolichopodidae s. lat., Microphorinae). Afr. Invertebr. 47: 335-346
  17. SINCLAIR B.J. & CUMMING J.M. 2006: The morphology, higherlevel phylogeny and classification of the Empidoidea (Diptera). Zootaxa 1180: 1-172
  18. SINCLAIR B.J. & SAIGUSA T. 2001: Revision of the world species of Ragas Walker (Diptera: Empidoidea). Entomol. Sci. 4: 507-522
  19. STUCKENBERG B.R. 1999: Antennal evolution in the Brachycera (Diptera), with a reassessment of terminology relating to the flagellum. Stud. Dipterol. 6: 33-48
  20. TUOMIKOSKI R. 1958: Mitteilungen ueber die Empididen (Dipt.) Finnlands. V. Die Gattung Iteaphila Zett. s. str. Ann. Entomol. Fenn. 24: 125-131